What they do is have the sound turned off when recording.That way the animal trainers can give the animals commands. then later they add the sounds. I seen that on extras of a disney film,I think it was old yeller.Now there is probably alot more computer generated stuff, but they still do it the way I described as well.I love the wimpering and child like noises.
I found it irratating, the first time I saw the movie and this time. It gives the animals a humanistic quality. The only thing they left out was the cub saying MAMMA.I love the wimpering and child like noises